The words on the left and right clue Latin bases, which can be combined to form full English words with Latin origins. The letters on the lines connecting the two pairs can be rearranged to spell out the full words, minus one letter.
- Porto (Carry) + Trans (Across) > Transporto = TRANSPOR[T]
- Ad (To) + Nihil (Nothing) > Annihilo = ANNI[H]ILATE
- In (Not) + Numerabilis (Countable) > Innumerablilis = INNUME[R]ABLE
- Capio (To take) + Pars (Part) > Participare = PARTICIPAT[E]
- Paene (Almost) + Umbra (Shadow) = P[E]NUMBRA
- Ambo (Both) + Dexter (Right) = AMBIDEX[T]ROUS
- Per (Through) + Tussis (Cough) = P[E]RTUSSIS
- Marina (Sea) + Aqua (Water) = Aqua Marina = AQUAMARIN[E]
- Nomen (Name) + Calare (Call) > Nomenclatura = NOMENCLA[T]URE
- Herba (Plant) + Cidium (Killing) = [H]ERBICIDE
In order of the left column, the missing letters spell out THREE TEETH. The word which comes from TRI (three) and DENS (teeth) is TRIDENT.